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Tetrapod Zoology

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Naish_profile_70_px.jpg With six years of phd work on theropod dinosaurs behind him, Darren Naish mostly spends long hours in the library, hunched over his laptop. But he gets out sometimes, and picks up litter and pursues exotic lizards across the British countryside, aiming all the while to publish his technical work on obscure Cretaceous dinosaurs. He also messes around with pterosaurs, swimming giraffes, British big cats and stuff like that. He has given up on the stupid idea of being a dedicated academic and ekes out a living as a technical consultant, editor and author. He can be contacted intermittently at eotyrannus (at) gmail dot com. For more biographical info go here. PLEASE NOTE: I am now completely unable to keep up with email correspondence, due to sheer volume and to time constraints. I do my best to respond to all queries and requests, but please don't be offended if I fail to reply.

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« Please name my flightless pterosaur | Main | Come back Lank, (nearly) all is forgiven »

Shemhazai and other flightless pterosaurs

Posted on: September 20, 2008 7:37 PM, by Darren Naish

how_azhdarchids_evolved_flightlessness.jpg

Sorry, another teaser - I haven't yet had time to post the full article (am aiming to do this on Monday). Again, all will be explained as goes the above [incoporating artwork by Mark Witton and yours truly]. Many, many, many thanks to everyone who made the 'name my flightless pterosaur' experiment such a soaraway (ha ha) success - I really enjoyed seeing your suggested names, many of which were excellent and/or very amusing. And the winner is...

Well, in the end I combined two different suggestions: the binomial is Shemhazai ptychocheirus. I don't expect you to agree with this choice, but I like the sound of it. If it needs correcting for etymological reasons let me know. Shemhazai was proposed by J. S. Lopes of Dinosauria Brasilis, while ptychocheirus (= 'folded hand') was from Tilsim. I was very fond of quite a few other names, including Nick Pharris's Pelargotitan altigradiens, Terradactylus telcontar from Adam Yates and Jenny Islander, and of course Darrendactylus naishi (albeit etymologically inappropriate), but in the end there can be only one. And you will notice that there is now another flightless azhdarchid... I suppose it also needs a name. Incidentally, I was going with Mike Keesey's Terrambulator skeksis until I googled Terrambulator and discovered that the term has already been used elsewhere (though not for an organism). Once again, thank you all.

DEFINITELY coming next: Come back Lank, all is forgiven (yes, it's about hypothetical flightless pterosaurs).

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Comments

1

Just out of interest... Are you planning on using Shemhazai ptychocheirus as an official name? Are there issues with the name now having been published without a formal description (i.e., does its appearance on this blog now make it a nomen nudum)?

I have very little knowledge of nomenclature so I'm asking not to be picky, but because I genuinely don't know.

Stewart

Posted by: Stewart Macdonald | September 20, 2008 8:41 PM

2

I imagine the animal is hypothetical (isn't it?)

Hmmm, for this new one ... Gigarhynchos camelopardaloides.

Posted by: William Miller | September 20, 2008 9:35 PM

3

Altodraco giraffasimilis. It means giraffe-like tall-dragon. At least I think it means that...

Hey, sorry for asking again, but now I'm curious: how can I say flitghless in latin, avolans???

Avolanodraco... Sounds pretty nice...

Posted by: Blue Frackle | September 20, 2008 10:05 PM

4

Another one??
This one seems to be more cursorial, with longer legs...
perhaps Dromopteron sp. or Dromodactylus sp.?
dromo-runner

(By the way, how do you make words in italic??)

Posted by: Frank | September 20, 2008 11:02 PM

5

Well, OK, I'll take a stab...

How about Ramphastops macrosceles (Gk) 'long-legged puffin-face'?

Posted by: Dr. Nick | September 21, 2008 1:53 AM

6

Argh. Fingers moving faster than brain. That should have been 'long-legged toucan-face'.

Posted by: Dr. Nick | September 21, 2008 1:54 AM

7

Ooh another one...

Dracopelargos terraphilus (Ground loving dragon stork)

Qilin cervasceles (Deer-legs qilin)

Arvocursor camelimimus (Camel-like plains-walker)

For the record Qilin is a giraffe-like mythical Chinese chimera said to bring good luck.

Posted by: Alec T | September 21, 2008 3:43 AM

8

Oh, Geezus. We're not...we're not validating Dixon, are we? Look at that giraffopterosaur. *weeps*

Posted by: Zach Miller | September 21, 2008 3:50 AM

9

Ah, the old, straight-necked, ghastly-winged, paper-thin skulled Quetzalcoatlus from Flickr makes a reappearance. That thing must be nearly 3 years old by now: might explain why I'm really not very keen on it anymore.

Funnily enough Darren, I was going to talk to you about speculative grounded pterosaurs in the near future. I've been thinking about what some of the more robustly jawed, massively built azhdarchids may have got up to if someone chopped their wings off. Oh, and tapejarids and dimorphodontids, too. Still, they'll have to wait: the real pterosaurs that form the basis of my thesis need more attention first.

Posted by: Mark Witton | September 21, 2008 4:35 AM

10

My thoughts are going out to whomever has to "design" the terrestrial anatomy of the arm. The hip and leg is noot really much of an issue, and a reflexibilization of the dorsals are not too problematic, as they should also lengthen a big, but I would think the arm much undergo a massive development alteration to be not only parasagitally but with the humerus more vertical, the distal end must twist about the axis, and rotate in position in toto for the purpose of bring the arm into stride alignment. Then there's the wrist, and the manus, which must slightly reorganize to be vertically load-bearing over transversely loadbearing horizontally. The arm was not designed for standing or walking, but exapted this way, contra the giraffid condition. This should be fascinating! (

Posted by: Jaime A. Headden | September 21, 2008 5:14 AM

11

I have no education in Greek or Latin, so these root-words are probably strung together wrong, but:

Dactylapteryx giraffamimus

"Finger-but-without-wing, giraffe-mimic"

Antaeodactyl alciorhynchus

Antaeus was a mythical Greek giant who had to touch the ground at all times or lose his strength so effectively that means:

"Land-bound-giant finger, auk-billed"

(I'm not at all sure how to put "alcis + rhynchus" together so the species name might be ungrammatical.)

The "dactyl" is not so meaningful with this animal, but I put it in there to hearken back to its pterosaur ancestors.

Feel free to mix-and-match each genus name with the other's species name, BTW.

Posted by: Stevo Darkly | September 21, 2008 5:33 AM

12

Oh, when I chose "auk-billed" for the species name above, I really wanted "puffin-billed" but could not find a meaningful Greek or Latin name for "puffin." (The puffin's genus name, Fratercula means "little brother" and I guess is nicely descriptive of puffins looking like little monks, but totally misleading here.)

And I had one more entry:

Cadopteryx parungulata

Which is as close as I can get to "fallen wing, sort-of-like-a-hoofed-mammal." (I just found a list of Latin words laiming that "cado" is the Latin word for "fallen to the ground.")

Posted by: Stevo Darkly | September 21, 2008 5:43 AM

13

Oops. I meant: I just found a list of Latin words CLAIMING that "cado" is the Latin word for "fallen to the ground."

Posted by: Stevo Darkly | September 21, 2008 5:59 AM

14
By the way, how do you make words in italic??

<i>This way.</> By using HTML.

The puffin's genus name, Fratercula means "little brother"

The funny thing is that this word is, in sum, feminine. No idea who came up with the -a at the end.

Posted by: David Marjanović | September 21, 2008 6:04 AM

15

<i>Oops.</i>

I just found a list of Latin words CLAIMING that "cado" is the Latin word for "fallen to the ground."

No. It means "I fall".

Posted by: David Marjanović | September 21, 2008 6:05 AM

16

David - Thanks, that was abundantly unclear from several different sources online. I guess the root word is "cadere" then? Would you know the proper form to stick in front of "pteryx" by chance?

And should my "parungulata" have been "parungulatus," I wonder?

Is there a doctor of classical languages in the house?

Posted by: Stevo Darkly | September 21, 2008 6:37 AM

17

I'm not a doctor of classical languages, but pteryx is feminine, so parungulata is correct.

Posted by: Andreas Johansson | September 21, 2008 7:01 AM

18

Darrendemon diatryma
Diatrymapteryx naishi

BTW, running pterosaurs are problematic from ecological point of view. This paper-thin, pneumatized skeleton stands no chance in competition with terrestrial runners and predators. Possible is evolution on some isolated island without saurischian predators, but, being good flyers and soarers, azhdarchids were unlikely to be stranded for a time needed to develop flightless species. In this way there is no flightless storks or raptors.

More likely are flightless tapejarids or other forms which are terrestrial and relatively weak flyers. Or maybe swimming pterosaurs turning into penguins or great auk equivalent.

BTW - I would look fr indication that some known pterosaurs could dive using wings, like modern auks.

Posted by: Jerzy | September 21, 2008 7:12 AM

19

This made my Sunday morning coffee tast even better than usual! Thanks Darren, also on behalf of the good old Greeks. Sadly unequipped to discuss pteroid function and postzygapophyses, I'm pleased to at least contribute in this modest way.

For this one I'd like to suggest Doradosaura 'gazelle lizard' (feminine is not necessary but sounds nice to me as doras is feminine; those who object to Maiasaura won't like this one either) or Pyrgosaurus 'tower lizard'.

Posted by: Tilsim | September 21, 2008 7:18 AM

20

Son of Clyde? Awesomeness defined. But perhaps forgotten:

Featuring sharp folding finger switchblades, with slashing claw. It seems a bit hard to see above, but a real terror in the hood, the wood, and even the water.

Posted by: DDeden | September 21, 2008 12:48 PM

21

Hey, just because it's the name of a machine doesn't mean it's unavailable! I WUZ ROBBED!!!

(Actually, I do like Shemhazai. And somebody has to use Tash inexorabilis for a pan-avian someday.)

Posted by: Mike Keesey | September 21, 2008 1:43 PM

22

WOW! I'm living my short moment of glory! Have my name chosen!

Beside Palaeontology,mythology and fallen angels was another of my favourite matters, since I was one of the creators of the Brazilian RPG "Ascensao e Queda" [Ascension and Fall], whose characters were children of fallen angels, in fact "de-fallen" angels who scaped from Hell during the Tunguska 1908's Explosion. More details in http://universogerminante.blogspot.com (Portuguese only)

SHEMHAZAI was the leader of Rebel Angels in Book of Enoch.

Posted by: J.S. Lopes | September 21, 2008 1:48 PM

23

Just got back from an excellent day at the zoo, during which I engaged in some surreal personal interaction with a ground hornbill. I will elaborate another time.

Want to respond briefly to one thing. Jerzy wrote...

BTW, running pterosaurs are problematic from ecological point of view. This paper-thin, pneumatized skeleton stands no chance in competition with terrestrial runners and predators. Possible is evolution on some isolated island without saurischian predators, but, being good flyers and soarers, azhdarchids were unlikely to be stranded for a time needed to develop flightless species.

I perhaps haven't explained it properly, but the idea here is that flightless azhdarchids evolved on predator-free islands (I think I stated this in the 'name my flightless pterosaur' article). However, I take issue with your 'paper-thin, pneumatized skeleton' comment. Yes, pterosaur bones were very thin (less than 1 mm cortical shaft thickness). But with a micro-layered, plywood-like cortical anatomy, trabecular struts, spiralling ridges on the internal cortical surfaces and other strengthening features, their bones were tremendously strong: in fact, one azhdarchid bone bitten by a theropod caused the theropod tooth tip to break off. Furthermore, work on limb bone strength and loading etc. shows that pterosaur limb bones were easily strong enough for running, standing launches, and for resisting the forces incurred by substantial activity. I would therefore disagree with the statement that hypothetical flightless pterosaurs might perform poorly against 'terrestrial runners and predators' for fundamental mechanical/ecological reasons.

In this way there is no flightless storks or raptors.

But there is no mechanical or ecological reason why there couldn't be flightless storks or raptors, it's that it never happened because it never happened, not because it couldn't. We know that it occurred among analogues: cranes became flightless at least once, and owls reduced or lost flight on several occasions.

BTW - I would look fr indication that some known pterosaurs could dive using wings, like modern auks.

There are no strong swimming or diving adaptations in known pterosaurs: we do not have any forms that might have behaved like auks, diving ducks or grebes. So, no!

Many thanks for your thoughts however, and to everyone else for theirs too. Commiserations to Mike Keesey!

Posted by: Darren Naish | September 21, 2008 2:38 PM

24
I guess the root word is "cadere" then?

That's the infinitive, yes.

Would you know the proper form to stick in front of "pteryx" by chance?

The stem is cad-, and then you need a connecting vowel, -o-. In other words, you were completely right, even though for the wrong reasons.

Posted by: David Marjanović | September 21, 2008 3:38 PM

25

Though... you want "fallen wing", not "fall wing". Try Casopteryx, assuming I'm not too tired to remember that correctly.

Posted by: David Marjanović | September 21, 2008 4:02 PM

26

Thanks for info on swimming pterosaurs.

So, where to look for fossils of this flightless pterosaur? What oceanic islands existed in Mesosoic?

Posted by: Jerzy | September 21, 2008 5:55 PM

27
So, where to look for fossils of this flightless pterosaur? What oceanic islands existed in Mesosoic?

Just one possibility among many...

Stilwell, J. D., Consoli, C. P., Sutherland, R., Salisbury, S., Rich, T. H., Vickers-Rich, P. A., Currie, P. J. & Wilson, G. J. 2006. Dinosaur sanctuary on the Chatham Islands, southwest Pacific: first record of theropods from the K-T boundary Takatika Grit. Palaeogeography, Palaeoclimatology, Palaeoecology 230, 243-250.

Posted by: Darren Naish | September 21, 2008 7:04 PM

28

the correct word for hand is "kheir", not "kheiros". It should be Shemhazai ptychocheir.

Posted by: J.S. Lopes | September 21, 2008 8:43 PM

29

J.S. Lopes wrote:

the correct word for hand is "kheir", not "kheiros". It should be Shemhazai ptychocheir.

True, but ptychocheirus is OK; it's an adjective meaning 'having folded hands'.

Deinonychus, among others, shows the same structure.

Posted by: Dr. Nick | September 21, 2008 9:44 PM

30

Andreas Johansson and David Marjanović, thank you for improving my language.

So, probably, my third entry is:

Casopteryx parungulata

("fallen wing, like an ungulate")

Posted by: Stevo Darkly | September 22, 2008 2:51 AM

31

Thanks, David Marjanović.

Posted by: Frank | September 22, 2008 11:06 AM

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