Human driven elephant evolution?

Why elephants are not so long in the tusk:

The average tusk size of African elephants has halved since the mid-19th century. A similar effect has been spotted in the Asian elephant population in India.

Researchers say it is an example of Darwinism in action, caused by the mass slaughter of dominant male elephants - but whereas evolution normally takes place over thousands of years, these changes have occurred within 150 years.

Zoologists at Oxford University fear that poaching and hunting of the largest male elephants, which also have the largest tusks, has changed the natural breeding behaviour of these animals. Their research has shown that the hunting of these large males for their ivory allows smaller males with shorter tusks to produce more calves. Over time the average tusk size decreases.

Of course it's wrong that "evolution normally takes place over thousands of years." Evolution is taking place now in many species, including the human species. Evolution most reductionistically understood is simply the change in allele frequencies over time. This can happen for a variety of reasons. Neutral theory arose to account for the stochastic changes in allele frequencies over time, especially discernible on the molecular level of DNA sequences. On the other hand, the process described above seems to be a rather standard account of evolution occurring via selection. Charles Darwin's original model of natural selection was strongly shaped by analogy to artificial selection imposed by humans upon domesticates; human predation has likely been a far stronger long term factor on the shape of faunal diversity the world over. In any case, one assumes that around 5 generations have elephants have lived over the past 150 years. One may also assume that tusk size is a quantitative character, influenced by many genes of small affect. Therefore, a model based on the breeder's equation seems plausible:

Response ~ Heritability X Selection Differential

The culling of elephant stocks strikes me as rather similar to truncation selection. A more interesting question to me is how often similar dynamics have been operative among other extant megafaunal species which have managed to survive and not been driven to extinction (yet). Finally, do note that if the change in tusk size is a function of quantitative selection, relaxation of selection (culling through poaching) should result in a bounce back of the trait value to its natural optimum.

Tags

More like this

I've talked about "the breeder's equation," R = h2S, before. R = response S = selection differential h2 = narrow sense heritability For example, if you have a population where the mean phenotypic value is 100, and you select a subpopulation with a mean value of 125 to breed the next generation,…
It's been nearly a month since I last posted on The Genetical Theory of Natural Selection. I've been holding off because I didn't know how to approach chapter 2, in many ways it is the most important and ambitious chapter (though not technically the most taxing). I think I will likely post twice…
Domestication has a long history. It predates the invention of writing by thousands of years. In the history of biology the study of domestication is closely connected to the emergence of experimental and theoretical biology out of the shadow of natural history. Chapter I of The Origin of Species…
Simpler mode of inheritance of transcriptional variation in male Drosophila melanogaster: Sexual selection drives faster evolution in males. The X chromosome is potentially an important target for sexual selection, because hemizygosity in males permits accumulation of alleles, causing tradeoffs in…

In layman's terms, evolution hasn't happened if you can't see it, i.e. if there is no a visible difference. In that sense evolution of large animals takes a long time.

The reduction of average size could also be a consequence of age distributions. If the current animals have few old individuals, all tusks haven't had enough time to grow to their full potential yet. How long does it take, anyway?

By Lassi Hippeläinen (not verified) on 22 Jan 2008 #permalink

In the course of not too long most mammals larger than a deer, especially carnivores, will no longer be viable except as protected species -- in many cases, only in zoos and wildlife parks. (That's already true of the cheetah and several species of tiger and rhino).

So the future evolution of these animals will be shaped by human culture, and they'll become pet species. They won't be selectively bred like dogs and horses were, I hope, but their formative environment will be human culture.

By John Emerson (not verified) on 22 Jan 2008 #permalink

I wonder if the same process created "polled" cattle in Scotland - Aberdeen Angus and other breeds - as people wanted drinking horns?!

Seems much more likely that people enjoyed not having to worry about getting gored and mated those animals preferentially. Since the fate of virtually all domestic cattle is to get dismantled at one point or another, people would get the horns one way or another--it's not like people are hunting cattle to get drinking horns (nor does it seem likely this was ever the case...it seems like the main benefit for killing a cow would always have been the vast chunk of meat it earned you.)

But who knows.

By Dr. Octoploid (not verified) on 22 Jan 2008 #permalink