The answers we seek: on 'goodbyes', the necks of caseids, and weird mystery sauropods

i-3232781e70c9bb9ac3070afb612de551-Ennatosaurus trio MS.jpg

Long-time blog readers will know that I am atrocious at keeping promises. And I will confess that part of the reason for titling an article 'Goodbye Tetrapod Zoology' was to cause a burst of panic, a rash of visitors (the strategy didn't really work: look at the counter... no spike on the graph). In seriousness, fear ye not oh followers, as I will indeed keep the blog ticking over, it's just that the only things I'll post will be short and sweet. And, unfortunately/fortunately, some bits of news come in that just demand a quick write-up...

Lurking in the Tet Zoo shadows are a number of professional palaeontologists, zoologists, artists and other workers whom many of you would be thrilled to be rubbing proverbial shoulders with. Obviously I'm thrilled (and sometimes a little terrified) to see that I'm attracting such people. I won't name names, but on this occasion I have cause to: Mike Skrepnick (website: Dinosaurs in Art) is a regular visitor, and after the brief post on caseids he has provided both images and some awesome information. In the caseids article, I included a photo of the late-surviving Russian taxon Ennatosaurus. It turns out that Mike had reason to produce several life restorations of Ennatosaurus when he worked with Hilary Madden (based at Robert Reisz's lab: she worked on Ennatosaurus for her master's thesis). With Mike's permission, I'm reproducing those images here: this is a world first, and I'm very proud to showcase entirely new work depicting such rarely-illustrated creatures (when writing the caseid post I was tempted to finish it with another of my laments about the lack of good artwork, but I've made this point enough times already, what with the aetosaur and temnospondyl articles, and don't want to become tedious). The images are © Mike Skrepnick, all rights reserved, used with permission.

i-0c45874a9ad2fb3138bd9edc88515c97-Caseid Digging MS.jpg

The illustration shown at the top of the article depicts a group of Ennatosaurus on a Permian hillside; they are accompanied by the low, scrubby cordiate Rufloria and the conifer Walchia. The Ennatosaurus skeleton that I showed previously (it's about 40 cm long I think) is apparently of a juvenile, and adults may have approached the giants like Cotylorhynchus in size. Big caseids like Ennatosaurus were living alongside biarmosuchians like Phthinosuchus and big dinocephalians like the predatory brithopodids Brithopus and Deuterosaurus and thick-skulled herbivorous tapinocephalids like the gigantic Ulemosaurus (Kemp 1982). Dicynodonts were doing well in Europe and southern Africa, but were rare in Russia at this time (King 1990). Anyway, some of these 'new' synapsids were big, reaching and exceeding 4 m in cases, so caseids had now been joined by other, more recently evolved giants.

i-2770ec77745e6db71c10a7832474f32c-Caseid Locomotion MS.jpg

As you can see from the black-and-white Skrepnick picture above, Mike confirms that caseids were likely to have been specialised diggers. Those of you wondering how the animal that owned the Oklahoma Cotylorhynchus skeleton (go here and here) might have reached the ground with its mouth will be interested to know that Madden and Reisz feel that the shoulder girdle in that specimen (and others) has been wrongly positioned, and needs to be raised so that the shoulder glenoid is higher. Furthermore, the head and neck needs to be repositioned closer to the ground. A more accurate depiction of the life posture is shown in the adjacent picture supplied by Mike (© Mike Skrepnick, all rights reserved, used with permission). The animals make more sense this way - but of course they still look plenty weird.

i-8fb165a941fa2bb93b2726657ccf19bf-Huanghetitan comp.jpg

So there you have it. Thanks again Mike for all this - very much appreciated. Finally (before I get back to the day job), what was that weird sauropod? Yes, it was a sauropod and not a Mesozoic dead-end roller-coster. Supplied by David Hone (of Ask A Biologist fame: remember to visit and ask any question you like... within reason), it depicts the recently unveiled skeleton of the titanosauriform Huanghetitan ruyangensis from the Upper Cretaceous Mangchuan Formation of Henan Province, China. Huanghetitan was described in 2006, with the type species being H. liujiaxiaensis (that's a lot of vowels), and in 2007 the type species and the new one H. ruyangensis were awarded their own 'family', Huanghetitanidae (Lü et al. 2007). With ribs at least 2.93 m long, H. ruyangensis is apparently the dinosaur with the 'deepest known body cavity from the Cretaceous of Asia'*. H. ruyangensis is known from sacral vertebrae, part of the tail, some ribs and part of the pelvis, so the neck and head you see here are reconstructions based on those of other sauropods. If Huanghetitan really is a basal titanosauriform, you might predict that it would have had a shorter, deeper more brachiosaur-like skull and far longer and much more gracile cervical ribs that overlap with one another, but what the hey.

* This prompts the tongue-in-cheek question: ok, now we know where the body cavity was from, but where was the dinosaur from? That's not the first time that wording like this has been used: the therizinosauroid Beipiaosaurus was published as 'A therizinosauroid dinosaur with integumentary structures from China' (Xu et al. 1999). So, ok, the integumentary structures were from China, but where was the dinosaur from?

So, as you can see, my 'goodbye' was perhaps somewhat over-rated. As I've said, I will keep the blog ticking over, but don't expect longish articles, like this one. Unless anything desperately cool or ugent comes along. Which it will.

Refs - -

Lü, J. Xu, L., Zhang, X. & Hu, W. 2007. A new gigantic sauropod dinosaur with the deepest known body cavity from the Cretaceous of Asia. Acta Geologica Sinica 81, 167-176.

Kemp, T. S. 1982. Mammal-like Reptiles and the Origin of Mammals. Academic Press (London).

King, G. 1990. The Dicynodonts: A Study in Palaeobiology. Chapman & Hall (London, New York).

Xu, X., Tang, Z.-l. & Wang, X.-l. 1999. A therizinosauroid dinosaur with integumentary structures from China. Nature 399, 350-354.


More like this

Among the many, many groups I have yet to cover on Tet Zoo are stem-group synapsids: Synapsida is the tetrapod clade that includes mammals and all of their relatives, and there is a long tradition of referring to non-mammalian synapsids as 'mammal-like reptiles' (other names include protomammals…
And so, here we are, at the end of it all. Ankylosaur week has come and gone, but oh what a week it was. As I said at the beginning, the whole aim was to save myself work and time by not producing anything new - and this worked, more or less. Did I clear the backlog? Did I hell, but at least I…
By now most people know that feathers are no longer unique to birds. Thanks mostly to a series of wonderful fossils from the Lower Cretaceous Yixian Formation of Liaoning Province, China, we now know that feathers first appeared in non-avian theropods, and were - later on - inherited by early…
First of all, many many thanks to everyone (both here and at SV-POW!) for the congrats regarding baby Emma. I am, shall we say, a little tired right now, but as you'll know if you've visited SV-POW! this morning, the good news as goes blogging is that I've been able to get lots of stuff prepared…

Darren, thanks again for the AAB plug!

For anyone who is interested, I took that photo at the recent press release in Zhengzhou, China. It did (kind of) make the news around the world (I spotted it on Yahoo, Washinton Post and a few others) as it turns out the locals have been eating their way quite happily through the 'dragon bones' in the hills for centuries. Who knows what has been lost.

On the upside there are tons of new animals out there (in addition to the one pictured), at least 1 other new sauropod, 2 or 3 ankylosaurs, a new dromaeosaur, oviraptorsaurid, 'carnosaur', numerous eggs and a new mammal. As ever with China we have high diversity, high numbers and high qualities of specimens. I am not exaggerating when I say this *could* be the next big fossil find. Some of the specimens are out of this world for quality and the size of the formation is huge and apparently, very productive.

All credit to Jungchang Lu for his work there in intial investigations and prompting the new wave of research which is just starting there.

There are more photos of the whole reconstruction online and videos etc. at:

I'll sign off now as this is supposed to be Darren's blog, not mine!

I'm glad to hear that TetZoo isn't really calling it quits! This blog is my primary window into the career in paleontology that almost was.

"In seriousness, fear ye not oh followers, as I will indeed keep the blog ticking over"

*sigh of relief* Good. The reason I did not comment earlier was that being told I'd have to go without Tet. Zoo. for a few weeks made me instantly go into withdrawl symptoms and pass out at the computer. Other than awakening to find the home row imprinted on my face, I'm fine, hah. Anyway, thanks for keeping things up despite the hectic schedule, Darren; it's impossible for me to read your posts and not learn something (in fact, many things) I hadn't previously been aware of.

Mike's pictures make all the difference in the world. Caseids are definatley bizarre, and his pictures (especially the last one) are beautiful!

"OK, his wooden leg was named Smith, but what did he call his other leg?"

Sad to say, I've yet to get a substantial, helpful answer from "Ask a Biologist".

By Nathan Myers (not verified) on 18 Jul 2007 #permalink

Humm, methinks I should have read this blog before posting on the previous one, I feel so foolish for attempting to name the above sauropod, ahhh well. I am glad I can still get my tetrapodzoology fix, life is good.

Excellent art by Mike. Beautiful drawing!

By Dan Varner (not verified) on 19 Jul 2007 #permalink

I'd just like to pile one more accolade onto Mike's drawings. I love the stride he depicted on that top creature.

Nice post Darren for one more time. So the animal in the sketches is Cotylorhynchus or Ennatosaurus?

By Georgios Georgalis (not verified) on 20 Jul 2007 #permalink

Great pictures!

Dicynodonts were doing well in Europe and southern Africa, but were rare in Russia at this time

That's because the Russian "Late Permian" is actually Middle Permian in its entirety (quoth A Geologic Time Scale 2004). This neatly explains why it's full of dinocephalians, too.

[from Darren: thanks for this. Why didn't anyone tell me sooner?]

By David Marjanović (not verified) on 29 Jul 2007 #permalink

Why didn't anyone tell me sooner?

Because nobody except my thesis supervisor has bought that huge book.

By David Marjanović (not verified) on 30 Jul 2007 #permalink